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For this purpose, ‘currentColor’ on a highlight pseudo-element’s ‘color’ property represents the ‘color’ of the next highlight pseudo-element layer below, falling back finally to the colors that would otherwise have been used (those applied by the originating element or an intervening pseudo-element such as ‘::first-line’ or ‘::first-letter’).
When we choose the “next highlight pseudo-element layer below”, do we consider only the layers that are actively highlighting the content in question? I think it makes sense to do so, in order for color:currentColor to effectively mean “highlighting with this pseudo doesn’t change the foreground color”. Otherwise, color:currentColor on ::selection always yields ::target-text’s default foreground color, even when that’s irrelevant to the content.
If so, what is the result of getComputedStyle(..., "::selection").color? The problem is that we need the resolved value, which would be indeterminate for a highlight with color:currentColor. Even if the active highlights were the same throughout the whole originating element, giving us a single unambiguous answer, we can only expose that answer if it didn’t come from ::spelling-error or ::grammar-error (#highlight-security).